The core hook of the series is its convoluted character relationships. The protagonist, Haruto, discovers that the girl he believed to be his older sister is actually his niece, and his first love—a woman from his past life—is reincarnated as his biological daughter. This "nested" complexity serves two purposes: it creates immediate comedic tension and sets a high-stakes emotional foundation. It challenges the reader to look past traditional labels of "sister" or "daughter" to see the soul of the individual beneath.
The story follows the life of Ritsu Onodera, a high school student who confesses his feelings to his crush, Masashi Otori. However, to his surprise, Otori reveals that he's not interested in Ritsu in a romantic way. Instead, Otori proposes a unusual arrangement: he'll pretend to be Ritsu's boyfriend in exchange for Ritsu helping him with his own unrequited love. Anehame Ore no Hatsukoi ga Jisshi na Wake ga Na...